Commencement Starts 11:30 Saturday
Due to rain in the forecast Saturday morning, the start time of the undergraduate ceremony has been delayed until 11:30 a.m. There will still be one ceremony outside in Biemesderfer Stadium. Gates open at 10 a.m.
Jean G. Boal, Ph.D
Dr. Jean Boal
Professor, Animal Behavior and Marine Biology.
Office: Roddy 283
M: 02:00 – 03:00 p.m.
T: 10:00 – 11:00 a.m.
R: 01:00 – 04:00 p.m.
B.S., Stanford University
Ph.D., University of North Carolina
BIOL 100 – General Biology
BIOL 281 – Behavioral Biology
BIOL 291 – Marine Biology
BIOL 390 – The Natural History of the Mid-Atlantic Shore
BIOL 471 – Topics in Biology
BIOL 472 – Seminar in Biology
BIOL 485/685 – Animal Behavior
Areas of Specialization:
Animal Behavior and Marine Biology.
I am interested in the evolution of cognitive behavior from a functional perspective. I am particularly interested in the relationship between an animal's learning abilities and its ecology and life history. My research has focused primarily on cephalopods (octopuses, cuttlefish, and squids) because they have large, complex brains compared to other invertebrates but we do not yet know why. Students participate actively in research with me here at Millersville University and at marine laboratories in Massachusetts and France. Prospective Students Please Note: Millersville University does not have an active graduate program in Biology at this time.
Selected Publication(s): (*student)
Jozet-Alves C, Darmaillacq A-S, Boal JG. In press. Navigation in cephalopods. In: Cephalopod Cognition. (A-S Darmaillacq, ed.)
Boal JG. 2011. Behavioral research methods for working with octopuses and cuttlefishes. Vie et Milieu 61: 203-210.
Walderon* MD, Nolt* K, Haas* RE, Prosser* KN, Holm* JB, Nagle GT, Boal JG. 2011. Distance chemoreception and the detection of conspecifics in Octopus bimaculoides. Journal of Molluscan Studies 77: 309-311. doi: 10.1093/mollus/eyr009.
Cummins SF, Boal JG, Buresch KC, Kuanpradit C, Sobhon P, Holm* JB, Degnan BM, Nagle GT, Hanlon RT. 2011. Extreme aggression in male squid induced by a ß-microseminoprotein-like pheromone. Current Biology 21(4): 322-327. doi: 10.1016/j.cub.2011.01.038.
Guibé* M, Boal JG, Dickel L. 2010. Early explosure to odors changes later visual prey preferences in cuttlefish. Journal of Developmental Psychobiology 1 Jun 2010 online in advance of print. DOI 10.1002/dev.20470.
Boal JG, Prosser* KN, Holm* JB, Simmons* TL, Haas* RE, Nagle GT. 2010. Sexually mature cuttlefish are attracted to the eggs of conspecifics. Journal of Chemical Ecology 36(8):834-836 DOI 10.1007/s10886-010-9816-0.
Alves* C, Boal JG, Dickel L. 2008. Short-distance navigation in cephalopods: a review. Cognitive Processing 9(4):239-247. DOI 10.1007/s10339-007-0192-9.
Hvorecny* LM, Grudowski* JL, Blakeslee* CJ, Simmons* TL, Roy* PR, Brooks* JA, Hanner* RM, Beigel* ME, Karson MA, Nichols* RH, Holm* JB, Boal JG. 2007. Octopuses (Octopus bimaculoides) and cuttlefishes (Sepia pharaonis, S. officinalis) can conditionally discriminate. Animal Cognition 10(4): 449-459. DOI 10.1007/s10071-007-0085-4.
Sigg* JE, Lloyd-Knight* KM, Boal JG. 2007. UV Light Influences covering behavior in the urchin Lytechinus variegatus. Journal of the Marine Biological Association 87: 1257-1261. DOI 10.1017/@0025315407055865.
Boal JG, Fenwick JW. 2007. Laterality in Octopus Eye Use? Animal Behaviour Forum 74(3):e1-e2. DOI.1016/j.anbehav.2006.12.026.
Komak*, S., L. Dickel, J.G. Boal, B.U. Budelmann. 2005. Behavioral responses of juvenile cuttlefish Sepia officinalis to local water movements. Marine and Freshwater Behaviour and Physiology 38(2): 117-125.
Boal, J.G., N. Shashar, M. Grable*, K. Vaughan*, E. Loew & R. Hanlon . 2004. Behavioral evidence for intraspecific signals with achromatic and polarized light by cuttlefish (Mollusca: Cephalopoda). Behaviour 141: 837-861.
Buresch, K.C., J.G. Boal, G.T. Nagle, J.Knowles*, R. Nobuhara, K. Sweeney* & R.T Hanlon. 2004. Experimental evidence that ovary and oviducal gland extracts influence male agonistic behavior in squids. Biological Bulletin 206: 1-3.
Buresch, K.C., J.G. Boal, J. Knowles*, J. DeBose*, A. Nichols*(MU), A. Erwin*, S.D. Painter, G.T. Nagle & R.T. Hanlon. 2003. Contact chemosensory cues in egg bundles elicit male-male agonistic conflicts in the squid Loligo pealei (Mollusca: Cephalopoda). Journal of Chemical Ecology. Vol 29(3): 547-560.
Boal, J. 1991. Complex learning in Octopus bimaculoides . American Malacological Bulletin 9(1):75-80.
Boal, J. 1980. Pacific harbor seal ( Phoca vitulina richardii). Haul out impact on the rocky midtidal zone. Marine Ecology Progress Series 2(4):265-269.